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protoconule

protoconule Zool.
  (-ˈkəʊnjʊl)
  [f. protocone + -ule.]
  An intermediate cusp between the protocone and the paracone of a mammalian upper molar tooth.

1888 [see protocone]. 1905 [see metaconule]. 1933 A. S. Romer Vertebr. Paleont. xii. 248 Between protocone and paracone there is often a smaller cusp, the protoconule. 1968 R. Zangerl tr. Peyer's Compar. Odontogr. 187 Intermediate cusps occurred along the trigon, a protoconule between protocone and paracone and a metaconule between protocone and metacone.

Oxford English Dictionary

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