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metaconule

metaconule Zool.
  (mɛtəˈkəʊnjʊl)
  [f. metacone + -ule.]
  An intermediate cusp between the hypocone and the metacone of a mammalian upper molar tooth.

1888 H. F. Osborn in Amer. Naturalist XXII. 1074 The Bunodont series are universally characterized by the initial or advanced development of the proto- and metaconules in the upper molars. 1905 Amer. Geologist XXXV. 244 The intermediate cusps (protoconule and metaconule) are both well-defined. 1968 R. Zangerl tr. Peyer's Compar. Odontol. 187 Intermediate cusps occurred.., a metaconule between protocone and metacone. 1971 W. D. Turnbull in A. A. Dahlberg Dental Morphol. & Evolution ix. 163 (caption) Specimen consists of the protocone and metaconules.

Oxford English Dictionary

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