Artificial intelligent assistant

How to quantitatively measure work done by a biological system? Thermodynamic efficiency can be expressed as the ratio of Work done(W) to Energy invested (Q). Thermodynamic efficiency= W/Q How can one measure work done by a biological system?

Measuring the work done by a biological system seems pretty impossible. Imagine how many different ways one cell of your body uses energy (ATP). You can't really measure all the work done by every cell on a macro scale. Metabolic efficiency has been defined as... "health". That seems just a little ambiguous. That's why we use things like averages to determine if energy use is normal or not, such as in metabolic age.

In short, work is a more tangible term in discrete physics examples, but there is so much complicated energy use in biological systems that total systemic work can't be easily defined.

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