This probably goes back to Lucas (1909), Adrian (1912), and Bernstein (1912). But the idea might have started from Helmholtz (1850).
* Lucas, K (1909). The ``all or none'' contraction of the amphibian skeletal muscle fibre. Journal of Physiology; 38: 113--133.
* Adrian, ED (1912). On the conduction of subnormal disturbances in normal nerve. Journal of Physiology; 45: 389--412
* Bernstein, J (1912). Electrobiologie. Braunschweig: Vieweg.
See also: Galvani's Spark: The Story of the Nerve Impulse and The great Era of English Electrophysiology: from Francis Gotch to Hodgkin and Huxley