Concerning < it was first used 1889 in the work _Arithmetices principia nova methodo exposita_ (page X):
> „Signum ϵ significat est. Ita a ϵ b legitur a est quoddam b“
This means something like (my translation which isn't the best!):
> The symbol ϵ means _is_. So a ϵ b has to be read as a _is a_ b
Here is a link to the work: <
The quoted sentence in the original work:
!reference