This is the same as or ("(while) never being satisfied"). This sense of is relatively rare in modern Japanese, but it's a traditional meaning of , and it should be listed in any monolingual dictionary. is a well-known set phrase.
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> ――
> **** ―
Note that is merely a temporary subject of the first half of the sentence including . The implicit main topic of this sentence is (humans), which is indicated in the previous sentence. can work as a noun ("hatred"), but in this context it's just 's continual form in its literal sense.
> () (Human's) selfishness surpasses the teaching of Amatsu; humans hate, deceit and repeat fighting, never being satisfied.