What does this Barbary Macaque facial expression mean? A girl pretended having a piece of food in her hand and pointed it to a Barbary Macaque in Gibraltar. As a response he made this facial expression to her. What does this mean? !GibraltarBarbaryMacaqueExpressions

It looks like it is trying too threat.

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Individual differences in scanpaths correspond with serotonin transporter genotype and behavioral phenotype in rhesus monkeys (Macaca mulatta)